The outcome of a 34-17 loss to the Denver Broncos was already pretty much decided at that point and frustration may have set in for a team skidding into its third consecutive defeat.
Boldin leads the team with nine penalties this season, including two unnecessary roughness infractions.
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Later, Williams tackled wide receiver Eric Decker well out of bounds after being beaten for a 13-yard catch.
"Anquan’s does not bother me at all," Harbaugh said. "Anquan was blocking back. It was a legal block. I saw it on tape. It was a block when Joe was running out and a guy was pursuing to tackle Joe, and it was a perfectly legal block.
"Cary’s was late. Cary’s was out of bounds and late. Cary would be the first to tell you that was a mistake.”